A youthful Liverpool side lost 2-1 at Union Saint-Gilloise in their final Europa League group game on Thursday, while Villarreal snatched a place in the last 16 from Rennes with a dramatic victory in France.

Bayer Leverkusen eased into the Europa League semi-finals after crushing hosts Union Saint-Gilloise of Belgium 4-1 on Thursday for a 5-2 aggregate win that stretched their unbeaten run to 12 straight matches across all competitions.